Turtles are reptiles of the order Testudines, characterised by a special bony or cartilaginous shell developed from their ribs that acts as a shield. “Turtle” may either refer to the Testudines as a whole, or to particular Testudines which make up a form taxon that is not monophyletic—see also sea turtle, terrapin, tortoise, and the discussion below. The order Testudines includes both extant and extinct species. The earliest known turtles date from 215 million years ago, making turtles one of the oldest reptile groups and a more ancient group than lizards, snakes and Crocodiles. Of the many species alive today some are highly endangered.